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The Pleistocene ( ; referred to colloquially as the ) is the geological epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated . Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding was regarded as being 1.806 million years (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the age used in . The name is a combination of () 'most' and (; as cænus) 'new'.

The and cooling trends of the preceding were continued in the Pleistocene. The climate was strongly variable depending on the glacial cycle, oscillating between cold and warmer , with the sea levels being up to lower than present at peak glaciation, allowing the periodic connection of Asia and North America via land bridge, and the covering of most of northern North America by the Laurentide Ice Sheet.

At the end of the preceding Pliocene, the previously isolated North and continents were joined by the Isthmus of Panama, causing a faunal interchange between the two regions and changing patterns, with the onset of glaciation in the Northern Hemisphere occurring around 2.7 million years ago. During the Early Pleistocene (2.58–0.8 Ma), of the genus originated in Africa and spread throughout . The end of the Early Pleistocene is marked by the Mid-Pleistocene Transition, with the cyclicity of glacial cycles changing from 41,000-year cycles to asymmetric 100,000-year cycles, making the climate variation more extreme. The witnessed the spread of modern humans outside Africa as well as the extinction of all other human species. Humans also spread to the Australian continent and the Americas for the first time.

The Late Pleistocene from around 50,000 years ago onwards was marked by the extinction of most large terrestrial animals outside of Africa, an extinction event otherwise unprecedented in the geological record due to its extreme size bias. Most authors suggest that climate change, the expansion of modern humans, or the combination of both were the likely causes of the extinctions.


Etymology
introduced the term "Pleistocene" in 1839 to describe strata in that had at least 70% of their molluscan fauna still living today. This distinguished it from the older , which Lyell had originally thought to be the youngest fossil rock layer. He constructed the name "Pleistocene" ('most new' or 'newest') from the πλεῖστος ( pleīstos) 'most' and καινός ( kainós ( as cænus) 'new'). From p. 621: "Toutefois, en même temps … et de substituer à la dénomination de Nouveau Pliocène celle plus abrégée de Pleistocène , tirée du grec pleiston, plus , et kainos, récent ." (However, at the same time that it became necessary to subdivide the two periods mentioned above, I found that the terms intended to designate these subdivisions were of an inconvenient length, and I have proposed to use in the future the word "Pliocene" for "old Pliocene", and to substitute for the name "new Pliocene" this shorter "Pleistocene", drawn from the Greek pleiston (most) and kainos (recent).) This contrasts with the immediately preceding ("newer", from πλείων ( pleíōn, "more") and kainós) and the immediately subsequent ("wholly new" or "entirely new", from ὅλος ( hólos, "whole") and kainós) epoch, which extends to the present time.


Dating
The Pleistocene has been dated from 2.580 million (±0.005) to 11,700 years BP with the end date expressed in as 10,000 carbon-14 years BP.For the top of the series, see:
(2025). 9780521781428, Cambridge University Press.
It covers most of the latest period of repeated , up to and including the cold spell. The end of the Younger Dryas has been dated to about 9700 BCE (11,700 calendar years BP). The end of the Younger Dryas is the official start of the current . Although it is considered an epoch, the Holocene is not significantly different from previous interglacial intervals within the Pleistocene.
(2025). 9780199913749, Oxford University Press. .
In the ICS timescale, the Pleistocene is divided into four stages or ages, the , Calabrian, (previously the unofficial "Middle Pleistocene"), and (unofficially the "Tarantian"). In addition to these international subdivisions, various regional subdivisions are often used.

In 2009 the International Union of Geological Sciences (IUGS) confirmed a change in time period for the Pleistocene, changing the start date from 1.806 to 2.588 million years BP, and accepted the base of the as the base of the Pleistocene, namely the base of the Monte San Nicola GSSP.Riccardi, Alberto C. (30 June 2009). "IUGS ratified ICS Recommendation on redefinition of Pleistocene and formal definition of base of Quaternary" International Union of Geological Sciences The start date has now been rounded down to 2.580 million years BP. The IUGS has yet to approve a , Global Boundary Stratotype Section and Point (GSSP), for the upper Pleistocene/Holocene boundary ( i.e. the upper boundary). The proposed section is the North Greenland Ice Core Project ice core 75° 06' N 42° 18' W. The lower boundary of the Pleistocene Series is formally defined magnetostratigraphically as the base of the Matuyama (C2r) , isotopic stage 103. Above this point there are notable extinctions of the calcareous nannofossils: .Gradstein, Felix M.; Ogg, James G. and Smith, A. Gilbert (eds.) (2005) A Geologic Time Scale 2004 Cambridge University Press, Cambridge, UK, p. 28, The Pleistocene covers the recent period of repeated glaciations.

The name has, in the past, been used to mean the last ice age. Formerly, the boundary between the two epochs was drawn at the time when the foraminiferal species Hyalinea baltica first appeared in the marine section at La Castella, Calabria, Italy.

(1975). 9780140551075, Penguin.
However, the revised definition of the , by pushing back the start date of the Pleistocene to 2.58 Ma, results in the inclusion of all the recent repeated glaciations within the Pleistocene.

Radiocarbon dating is considered to be inaccurate beyond around 50,000 years ago. Marine isotope stages (MIS) derived from Oxygen isotopes are often used for giving approximate dates.


Deposits
Pleistocene non-marine are found primarily in fluvial deposits, lakebeds, slope and deposits as well as in the large amounts of material moved about by glaciers. Less common are deposits, and volcanic deposits (lavas, ashes). Pleistocene marine deposits are found primarily in shallow marine basins mostly (but with important exceptions) in areas within a few tens of kilometres of the modern shoreline. In a few geologically active areas such as the Southern California coast, Pleistocene marine deposits may be found at elevations of several hundred metres.


Paleogeography and climate
The modern were essentially at their present positions during the Pleistocene, the upon which they sit probably having moved no more than relative to each other since the beginning of the period. In glacial periods, the sea level would drop by up to lower than today during peak glaciation, exposing large areas of the present continental shelf as dry land.

According to (through collected data), the Pleistocene's overall climate could be characterised as a continuous El Niño with in the south weakening or heading east, warm air rising near , warm water spreading from the west Pacific and the to the east Pacific, and other El Niño markers.National Geographic Channel, Six Degrees Could Change The World, Mark Lynas interview. Retrieved 14 February 2008.


Glacial features
Pleistocene climate was marked by repeated glacial cycles in which continental glaciers pushed to the 40th parallel in some places. It is estimated that, at maximum glacial extent, 30% of the Earth's surface was covered by ice. In addition, a zone of stretched southward from the edge of the glacial sheet, a few hundred kilometres in , and several hundred in . The mean annual temperature at the edge of the ice was ; at the edge of the permafrost, .

Each glacial advance tied up huge volumes of water in continental ice sheets thick, resulting in temporary sea-level drops of or more over the entire surface of the Earth. During interglacial times, such as at present, drowned coastlines were common, mitigated by isostatic or other emergent motion of some regions.

The effects of glaciation were global. was ice-bound throughout the Pleistocene as well as the preceding Pliocene. The were covered in the south by the ice cap. There were glaciers in and . The current decaying glaciers of , Mount Kilimanjaro, and the in east and central Africa were larger. Glaciers existed in the mountains of and to the west in the .

In the northern hemisphere, many glaciers fused into one. The Cordilleran Ice Sheet covered the North American northwest; the east was covered by the . The Fenno-Scandian ice sheet rested on , including much of Great Britain; the Alpine ice sheet on the . Scattered domes stretched across and the Arctic shelf. The northern seas were ice-covered.

South of the ice sheets large lakes accumulated because outlets were blocked and the cooler air slowed evaporation. When the Laurentide Ice Sheet retreated, north-central North America was completely covered by . Over a hundred basins, now dry or nearly so, were overflowing in the North American west. , for example, stood where Great Salt Lake now does. In Eurasia, large lakes developed as a result of the runoff from the glaciers. Rivers were larger, had a more copious flow, and were . African lakes were fuller, apparently from decreased evaporation. Deserts, on the other hand, were drier and more extensive. Rainfall was lower because of the decreases in oceanic and other evaporation.

It has been estimated that during the Pleistocene, the East Antarctic Ice Sheet thinned by at least 500 meters, and that thinning since the Last Glacial Maximum is less than 50 meters and probably started after c. 14 ka.


Major events
During the 2.5 million years of the Pleistocene, numerous cold phases called (Quaternary ice age), or significant advances of continental ice sheets, in Europe and North America, occurred at intervals of approximately 40,000 to 100,000 years. The long glacial periods were separated by more temperate and shorter which lasted about 10,000–15,000 years. The last cold episode of the last glacial period ended about 10,000 years ago. Over 11 major glacial events have been identified, as well as many minor glacial events. A major glacial event is a general glacial excursion, termed a "glacial". Glacials are separated by "interglacials". During a glacial, the glacier experiences minor advances and retreats. The minor excursion is a "stadial"; times between stadials are "interstadials".

These events are defined differently in different regions of the glacial range, which have their own glacial history depending on latitude, terrain and climate. There is a general correspondence between glacials in different regions. Investigators often interchange the names if the glacial geology of a region is in the process of being defined. However, it is generally incorrect to apply the name of a glacial in one region to another.

For most of the 20th century, only a few regions had been studied and the names were relatively few. Today the geologists of different nations are taking more of an interest in Pleistocene glaciology. As a consequence, the number of names is expanding rapidly and will continue to expand. Many of the advances and stadials remain unnamed. Also, the terrestrial evidence for some of them has been erased or obscured by larger ones, but evidence remains from the study of cyclical climate changes.

The glacials in the following tables show historical usages, are a simplification of a much more complex cycle of variation in climate and terrain, and are generally no longer used. The headings "Glacial 1" to "Glacial 4" are designations indicating the four most recent glacials, with "Glacial 4" being the most recent. These names have been abandoned in favour of numeric data because many of the correlations were found to be either inexact or incorrect and more than four major glacials have been recognised since the historical terminology was established. (contains a summary of how and why the Nebraskan, Aftonian, Kansan, and Yarmouthian stages were abandoned by modern stratigraphers).

+ Historical names of the "four major" glacials in four regions. ! Region ! Glacial 1 ! Glacial 2 ! Glacial 3 ! Glacial 4
AlpsMindelWürm
North Europe
British Isles
Midwest U.S.KansanIllinoianWisconsinan

+ Historical names of interglacials. ! Region ! Interglacial 1 ! Interglacial 2 ! Interglacial 3
Alps
North EuropeWaalianHolsteinian
British Isles
Midwest U.S.Yarmouthian

Corresponding to the terms glacial and interglacial, the terms pluvial and interpluvial are in use (Latin: pluvia, rain). A pluvial is a warmer period of increased rainfall; an interpluvial is of decreased rainfall. Formerly a pluvial was thought to correspond to a glacial in regions not iced, and in some cases it does. Rainfall is cyclical also. Pluvials and interpluvials are widespread.

There is no systematic correspondence between pluvials to glacials, however. Moreover, regional pluvials do not correspond to each other globally. For example, some have used the term "Riss pluvial" in Egyptian contexts. Any coincidence is an accident of regional factors. Only a few of the names for pluvials in restricted regions have been stratigraphically defined.


Palaeocycles
The sum of transient factors acting at the Earth's surface is cyclical: climate, ocean currents and other movements, wind currents, temperature, etc. The waveform response comes from the underlying cyclical motions of the planet, which eventually drag all the transients into harmony with them. The repeated glaciations of the Pleistocene were caused by the same factors.

The Mid-Pleistocene Transition, approximately one million years ago, saw a change from low-amplitude glacial cycles with a dominant periodicity of 41,000 years to asymmetric high-amplitude cycles dominated by a periodicity of 100,000 years.

However, a 2020 study concluded that ice age terminations might have been influenced by since the Mid-Pleistocene Transition, which caused stronger summers in the Northern Hemisphere.


Milankovitch cycles
Glaciation in the Pleistocene was a series of glacials and interglacials, stadials and interstadials, mirroring periodic climate changes. The main factor at work in climate cycling is now believed to be Milankovitch cycles. These are periodic variations in regional and planetary solar radiation reaching the Earth caused by several repeating changes in the Earth's motion. The effects of Milankovitch cycles were enhanced by various positive feedbacks related to increases in atmospheric carbon dioxide concentrations and Earth's .

Milankovitch cycles cannot be the sole factor responsible for the variations in climate since they explain neither the long-term cooling trend over the Plio-Pleistocene nor the millennial variations in the Greenland Ice Cores known as Dansgaard-Oeschger events and . Milankovitch pacing seems to best explain glaciation events with periodicity of 100,000, 40,000, and 20,000 years. Such a pattern seems to fit the information on climate change found in oxygen isotope cores.


Oxygen isotope ratio cycles
In oxygen isotope ratio analysis, variations in the ratio of to (two of ) by (measured by a mass spectrometer) present in the of oceanic is used as a diagnostic of ancient ocean temperature change and therefore of climate change. Cold oceans are richer in , which is included in the tests of the microorganisms () contributing the calcite.

A more recent version of the sampling process makes use of modern glacial ice cores. Although less rich in than seawater, the snow that fell on the glacier year by year nevertheless contained and in a ratio that depended on the mean annual temperature.

Temperature and climate change are cyclical when plotted on a graph of temperature versus time. Temperature coordinates are given in the form of a deviation from today's annual mean temperature, taken as zero. This sort of graph is based on another isotope ratio versus time. Ratios are converted to a percentage difference from the ratio found in standard mean ocean water (SMOW).

The graph in either form appears as a with . One half of a period is a Marine isotopic stage (MIS). It indicates a glacial (below zero) or an interglacial (above zero). Overtones are stadials or interstadials.

According to this evidence, Earth experienced 102 MIS stages beginning at about 2.588 in the Early Pleistocene . Early Pleistocene stages were shallow and frequent. The latest were the most intense and most widely spaced.

By convention, stages are numbered from the Holocene, which is MIS1. Glacials receive an even number and interglacials receive an odd number. The first major glacial was MIS2-4 at about 85–11 ka BP. The largest glacials were 2, 6, 12, and 16. The warmest interglacials were 1, 5, 9 and 11. For matching of MIS numbers to named stages, see under the articles for those names.


Fauna
Both marine and continental faunas were essentially modern but with many more large land mammals such as , , , , , , , , , species within and others. Isolated landmasses such as , , and islands in the Pacific saw the evolution of large birds and even reptiles such as the , , Haast's eagle, , and .

The severe climatic changes during the Ice Age had major impacts on the fauna and flora. With each advance of the ice, large areas of the continents became depopulated, and plants and animals retreating southwards in front of the advancing glacier faced tremendous stress. The most severe stress resulted from drastic climatic changes, reduced living space, and curtailed food supply. A major of large (), which included , , , , the woolly rhinoceros, various , such as the ; , , , , , , , and , began late in the Pleistocene and continued into the Holocene. also became extinct during this period. At the end of the last ice age, animals, smaller mammals like , migratory birds, and swifter animals like had replaced the megafauna and migrated north. Late Pleistocene were more slender and had longer legs than their descendants today. Scientists believe that the change in predator fauna after the late Pleistocene extinctions resulted in a change of body shape as the species adapted for increased power rather than speed.

The extinctions hardly affected Africa but were especially severe in where native and were wiped out.

  • Asian land mammal ages (ALMA) include , , and .
  • European land mammal ages (ELMA) include the , Galerian, and Aurelian
  • North American land mammal ages (NALMA) include (4.75–1.8), (1.8–0.24) and (0.24–0.01) in millions of years. The Blancan extends significantly back into the Pliocene.
  • South American land mammal ages (SALMA) include (2.5–1.5), (1.5–0.3) and (0.3–0.01) in millions of years. The Uquian previously extended significantly back into the Pliocene, although the new definition places it entirely within the Pleistocene.

In July 2018, a team of scientists in collaboration with Princeton University announced that they had brought two female frozen in , from around 42,000 years ago, back to life. The two nematodes, at the time, were the oldest confirmed living animals on the planet.

File:Ice age fauna of northern Spain - Mauricio Antón.jpg|Pleistocene of , including , cave lions eating a , , and woolly rhinoceros File:Pleistocene SA.jpg|Pleistocene of , including and two


Humans
The of anatomically modern humans took place during the Pleistocene. At the beginning of the Pleistocene species were still present, as well as early human ancestors, but during the lower Palaeolithic they disappeared, and the only species found in fossilic records is for much of the Pleistocene. appear along with Homo erectus, some 1.8 million years ago, replacing the more primitive industry used by Australopithecus garhi and by the earliest species of Homo. The Middle Paleolithic saw more varied speciation within Homo, including the appearance of about 300,000 years ago. Artifacts associated with modern human behavior are unambiguously attested starting 40,000–50,000 years ago.

According to mitochondrial timing techniques, modern humans migrated from Africa after the in the Middle Palaeolithic during the , spreading all over the ice-free world during the late Pleistocene.Stringer, C.B. (1992) "Evolution of early modern humans" In: Jones, Steve; Martin, R. and Pilbeam, David R. (eds.) (1992) The Cambridge encyclopedia of human evolution Cambridge University Press, Cambridge, , pp. 241–251. A 2005 study posits that humans in this migration interbred with forms already outside of Africa by the late Pleistocene, incorporating archaic human genetic material into the modern human gene pool.


See also


Explanatory notes


External links

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